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The cochlea and tonotopic organisation

Each point on the basilar membrane responds best to one particular frequency. The base (near the oval window) is narrow and stiff and responds to high frequencies. The apex (near the helicotrema) is wide and flexible and responds to low frequencies. Stiffness falls roughly a hundredfold from base to apex. (von Békésy, 1960; Greenwood, 1990)

The wave always starts at the base and travels to the place of its characteristic frequency, where it peaks and then dies away. A low-frequency sound therefore moves a much longer stretch of the membrane, while a high-frequency sound stays within the first few millimetres of the base.

Apex · 20 HzBase · 20 kHz

The sections in the right-hand panel let you explore types of hearing loss, a cochlear implant and a cross-section of the duct.

Drag to rotate; pinch or use the wheel to zoom. Tap a point on the membrane to hear that point's frequency. The space bar starts and stops the sound; the left and right arrows change the frequency by a third of an octave, the up and down arrows change the level by 5 dB.

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